You actually make the reservation with the rental company through a portal from the Costco website. Any selected discount codes are automatically entered into your reservation.
Alternatively, you can do what I do and make a note of the desired discount codes and then manually log into my account at the rental website, preserving any additional benefits I have there under my avvound (e.g. Avis Preferred/First, Budget Fastbreak, etc). I then manually enter the codes into a reservation request.
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